Б. Протондвижущая сила

Ионы гидроксония («Н+-ионы») также могут формировать электрохимический градиент. Такой протонный градиент имеет решающее значение для клеточного синтеза АТФ (см. Белки главного комплекса гисто-совместимости). Как и в случае других ионов, свободная энергия переноса протона (разность между электрохимическими потенциалами протонов на двух сторонах мембраны) зависит от градиента концентрации, то есть от разности pH (ΔрН) по ту и другую стороны мембраны. Кроме того, определённый вклад вносит и трансмембранный потенциал ΔΨ (см. А). Обе эти величины формируют протондвижущую силу Δp, являющуюся мерой работы ΔΨG, которую может совершать Н+-градиент. Таким образом, протонный градиент через внутреннюю митохондриальную мембрану (см. Синтез АТФ) даёт примерно 24 кДж на моль переносимых ионов Н+.


Метаболизм. Энергетика / Сохранение энергии на мембранах

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